Effects of oncogenic transformation on cell motility and the cytoskeleton; cell cycle checkpoints based on the cytoskeleton.
- 1982: MD, Johns Hopkins University, Baltimore
- 1983: PhD, cell biology, Johns Hopkins University
- 1983 - 1984: Postdoctoral fellow, cell biology, Johns Hopkins University, Baltimore
- 1984 - 1985: Intern, pathology, Washington University, St. Louis
- 1985 - 1987: Postdoctoral fellow, cell biology, Washington University
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