Timothy Ley, MD

Timothy Ley, MD

Primary Academic Title

Lewis T. and Rosalind B. Apple Professor of Oncology in Medicine, Division of Oncology, Section of Stem Cell Biology, Washington University School of Medicine

Research Interest

Regulation of globin gene expression; regulation of gene expression during myelomonocytic development (cathepsin G) and T-Cell activation (Gzm B and H); functions of serine proteases in immune effector cells.


  • 1978: MD, Washington University, St. Louis


  • 1978 - 1980: Intern and resident, Massachusetts General Hospital, Boston
  • 1980 - 1983: Postdoctoral fellow, molecular hematology, National Heart, Blood and Lung Institute, Bethesda, Md.
  • 1083 - 1984: Fellow, hematology/oncology, Washington University, St. Louis

Journal Articles

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