Regulation of globin gene expression; regulation of gene expression during myelomonocytic development (cathepsin G) and T-Cell activation (Gzm B and H); functions of serine proteases in immune effector cells.
- 1978: MD, Washington University, St. Louis
- 1978 - 1980: Intern and resident, Massachusetts General Hospital, Boston
- 1980 - 1983: Postdoctoral fellow, molecular hematology, National Heart, Blood and Lung Institute, Bethesda, Md.
- 1083 - 1984: Fellow, hematology/oncology, Washington University, St. Louis
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